Skills in Borderlands 4 are the backbone of any Vault Hunter build. They are grouped under distinctive Skill Trees. There are 3 Skill Trees per Vault Hunter, and each Tree has 3 Skill Tree Branches.

The majority of a Skill Tree consists of Passive Skills. They can be either Conditional - trigger whenever a Vault Hunter does something - or Passive - work all the time. Besides Passive Skills, each Tree has one Action Skill and a couple of Augments and Capstones. Vault Hunter's Action Skill, available from the start but needs to be assigned from a Skill Tree, is a powerful Ability they can use when not on cooldown. Augments provide various additional bonuses to an Action Skill. Lastly, Capstones are build-defining heavy modifications to Action Skills. Capstones are positioned at the end of each Skill Tree Branch.

Both Augments and Capstones don't need Skill Point investment in themselves to be unlocked. Instead, they become available once a certain total number of Skill Points invested in a Skill Tree is reached. In total, each Skill Tree offers 3 Capstones and 5 Augments. Please notice that a Vault Hunter can only equip one Augment and one Capstone in addition to an Action Skill. Augments and Capstones from a specific Skill Tree are tied to the Action Skill of that Skill Tree.

Vault Hunter Skill Trees

 

 

 Siren Skill Trees

The Fourth Seal
Vexcalation
Here Comes Trouble



dead ringer skill tree vex borderlands 4 wiki guide

Siren: The Fourth Seal 


Level 1 Skills

sanguine fiends vex skill borderlands 4 wiki guide
Sanguine Fiends


5 Points

idle hands vex skill borderlands 4 wiki guide
Idle Hands


3 Points
 

cold iron vex skill borderlands 4 wiki guide
Cold Iron


5 Points

Level 2 Skills

grave assault vex skill borderlands 4 wiki guide
Grave Assault


5 Points

ghast blaster vex skill borderlands 4 wiki guide
Ghast Blaster


Augment
 

coven vex skill borderlands 4 wiki guide
Coven


Augment
 

cursed blade vex skill borderlands 4 wiki guide
Cursed Blade


5 Points

Level 3 Skills

blood sacrifice vex skill borderlands 4 wiki guide
Blood Sacrifice


1 Point

grave fiends vex skill borderlands 4 wiki guide
Grave Fiends


5 Points
 

grave quickening vex skill borderlands 4 wiki guide
Grave Quickening


5 Points

extra cursed vex skill borderlands 4 wiki guide
Extra Cursed


1 Point

Left Branch Level 1 Skills

wither vex skill borderlands 4 wiki guide
Wither


Augment
 

prismatic ichor vex skill borderlands 4 wiki guide
Prismatic Ichor


5 Points

Left Branch Level 2 Skills

keen suffering vex skill borderlands 4 wiki guide
Keen Suffering


5 Points

ghasterbait vex skill borderlands 4 wiki guide
Ghasterbait


3 Points
 

shot to go vex skill borderlands 4 wiki guide
Shot to Go!


3 Points

Left Branch Level 3 Skills

necrosis vex skill borderlands 4 wiki guide
Necrosis


5 Points

heirloom vex skill borderlands 4 wiki guide
Heirloom


Capstone

Center Branch Level 1 Skills

hellish harvest vex skill borderlands 4 wiki guide
Hellish Harvest


Augment
 

recurrence vex skill borderlands 4 wiki guide
Recurrence


5 Points

Center Branch Level 2 Skills

reverburnations vex skill borderlands 4 wiki guide
Reverburnations


5 Points

haruspex vex skill borderlands 4 wiki guide
Haruspex


5 Points

Center Branch Level 3 Skills

burnt offering vex skill borderlands 4 wiki guide
Burnt Offering


5 Points

grim reaper vex skill borderlands 4 wiki guide
Grim Reaper


Capstone
 

pincushion vex skill borderlands 4 wiki guide
Pincushion


1 Point

Right Branch Level 1 Skills

banshees wail vex skill borderlands 4 wiki guide
Banshee's Wail


Augment
 

obedient fiends vex skill borderlands 4 wiki guide
Obedient Fiends


5 Points

Right Branch Level 2 Skills

spirit bomb vex skill borderlands 4 wiki guide
Spirit Bomb


3 Points

ancient rites vex skill borderlands 4 wiki guide
Ancient Rites


5 Points

 

Right Branch Level 3 Skills

diener vex skill borderlands 4 wiki guide
Diener


1 Point

dark pact vex skill borderlands 4 wiki guide
Dark Pact


Capstone

 

 

incarnate skill tree vex borderlands 4 wiki guide

Siren: Vexcalation 



Level 1 Skills

guns and needles vex skill borderlands 4 wiki guide
Guns And Needles


3 Points

volatile essence vex skill borderlands 4 wiki guide
Volatile Essence


5 Points
 

grave sights vex skill borderlands 4 wiki guide
Grave Sights


5 Points

Level 2 Skills

geistwave vex skill borderlands 4 wiki guide
Geistwave


Augment
 

bloodletter vex skill borderlands 4 wiki guide
Bloodletter


5 Points

energy vampire vex skill borderlands 4 wiki guide
Energy Vampire


Augment
 

grave implements vex skill borderlands 4 wiki guide
Grave Implements


5 Points

Level 3 Skills

channeling vex skill borderlands wiki guide
Channeling


5 Points
 

grave power vex skill borderlands 4 wiki guide
Grave Power


3 Points

target prescience vex skill borderlands 4 wiki guide
Target Prescience


5 Points

Left Branch Level 1 Skills

iron maiden vex skill borderlands 4 wiki guide
Iron Maiden


Augment
 

conjunction vex skill borderlands 4 wiki guide
Conjunction


5 Points

Left Branch Level 2 Skills

dreadlight vex skill borderlands 4 wiki guide
Dreadlight


5 Points
 

full moon vex skill borderlands 4 wiki guide
Full Moon


5 Points

Left Branch Level 3 Skills

havoc vex skill borderlands 4 wiki guide
Havoc


5 Points

desecration vex skill borderlands 4 wiki guide
Desecration


Capstone

Center Branch Level 1 Skills

drink full vex skill borderlands 4 wiki guide
Drink Full


5 Points
 

keen mind vex skill borderlands 4 wiki guide
Keen Mind


5 Points

Center Branch Level 2 Skills

phase pocket vex skill borderlands 4 wiki guide
Phase Pocket


3 Points

mortal terror vex skill borderlands 4 wiki guide
Mortal Terror


5 Points
 

grave elements vex skill borderlands 4 wiki guide
Grave Elements


5 Points

Center Branch Level 3 Skills

vibe check vex skill borderlands 4 wiki guide
Vibe Check


Capstone

strike me down borderlands 4 wiki guide
Strike Me Down


1 Point

Right Branch Level 1 Skills

grave vitality vex skill borderlands 4 wiki guide
Grave Vitality


5 Points

grave harvest vex skill borderlands 4 wiki guide
Grave Harvest


Augment
 

ars arcana vex skill borderlands 4 wiki guide
Ars Arcana


5 Points

Right Branch Level 2 Skills

a deeper grave vex skill borderlands 4 wiki guide
A Deeper Grave


3 Points

blood shot vex skill borderlands 4 wiki guide
Blood Shot


5 Points

 

essence leech vex skill borderlands 4 wiki guide
Essence Leech


5 Points

Right Branch Level 3 Skills

heartpiercer vex skill borderlands 4 wiki guide
Heartpiercer


Capstone

blood is power vex skill borderlands 4 wiki guide
Blood Is Power


3 Points

phase phamiliar skill tree vex borderlands 4 wiki guide

Siren: Here Comes Trouble 



Level 1 Skills

grave sustenance vex skill borderlands 4 wiki guide
Grave Sustenance


5 Points

enchantment vex skill borderlands 4 wiki guide
Enchantment


5 Points
 

blood is magic vex skill borderlands 4 wiki guide
Blood Is Magic


5 Points

hasty fiends vex skill borderlands 4 wiki guide
Hasty Fiends


3 Points

Level 2 Skills

grave command vex skill borderlands wiki guide
Grave Command


3 Points
 

overprotective vex skill borderlands 4 wiki guide
Overprotective


5 Points

blink strike vex skill borderlands 4 wiki guide
Blink Strike


Augment
 

yowl and bay vex skill borderlands 4 wiki guide
Yowl And Bay


5 Points

Level 3 Skills

mourning vex skill borderlands 4 wiki guide
Mourning


5 Points

claw and bang vex skill borderlands 4 wiki guide
Claw And Bang


5 Points
 

overheal vex skill borderlands 4 wiki guide
Overheal


3 Points

vorpal fiends vex skill borderlands 4 wiki guide
Vorpal Fiends


5 Points

Left Branch Level 1 Skills

icy attunement vex skill borderlands 4 wiki guide
Icy Attunement


5 Points

hexus nexus vex skill borderlands 4 wiki guide
Hexus Nexus


Augment
 

blast rites vex skill borderlands 4 wiki guide
Blast Rites


5 Points

Left Branch Level 2 Skills

overspirited vex skill borderlands 4 wiki guide
Overspirited


5 Points

blasted fiends borderlands 4 wiki guide
Blasted Fiends


5 Points
 

corporeal vex skill borderlands 4 wiki guide
Corporeal


3 Points

Left Branch Level 3 Skills

shiver vex skill borderlands 4 wiki guide
Shiver


5 Points

trouble bubble vex skill borderlands 4 wiki guide
Trouble Bubble


Capstone

Center Branch Level 1 Skills

vorpal fang vex skill borderlands 4 wiki guide
Vorpal Fang


Augment
 

keen fiends vex skill borderlands 4 wiki guide
Keen Fiends


5 points

Center Branch Level 2 Skills

infernal sum vex skill borderlands 4 wiki guide
Infernal Sum


5 Points
 

reawakening vex skill borderlands 4 wiki guide
Reawakening


5 Points

Center Branch Level 3 Skills

tail whip vex skill borderlands 4 wiki guide
Tail Whip


1 Point

double trouble vex skill borderlands 4 wiki guide
Double Trouble


Capstone

Right Branch Level 1 Skills

beast mode vex skill borderlands 4 wiki guide
Beast Mode


Augment
 

grave current vex skill borderlands 4 wiki guide
Grave Current


5 Points

Right Branch Level 2 Skills

deadwire vex skill borderlands 4 wiki guide
Deadwire


5 Points

grave thirst vex skill borderlands 4 wiki guide
Grave Thirst


5 Points

 

infusion vex skill borderlands 4 wiki guide
Infusion


5 Points

Right Branch Level 3 Skills

stormblessed vex skill borderlands 4 wiki guide
Stormblessed


5 Points

unsealed vex skill borderlands 4 wiki guide
Unsealed


Capstone

 




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